Diversity: Only three genera, with willows (Salix - ca. 400 species) and poplars (Populus - ca. 35 species) in the North American flora but, as was the case with taxa of the Hamamelidae, the relatively small number of species includes many with a wide distribution and, as a result, plants of the family - all woody (shrubs or trees) are conspicuous elements of a given flora and frequently encountered.
Distribution: Throughout temperate parts of the World with maximum diversity toward the north and a strong ecological preference toward habitats associated with water. In our local flora, taxa of both genera will be found beside ponds and along streams and rivers. The Texas flora includes 17 species and 6 infraspecific taxa.
Floral structure:

Significant features:
Floral structure of the Salicaceae is an expression of extreme
reduction.
The pistillate flower is represented by only a single, compound pistil
and the staminate flower is, in Salix species, just two
stamens. These imperfect flowers are produced in catkins
and most species of the family are dioecious.
While the reproductive adaptive syndrome of the Salicaceae mimics that
of the Hamamelidae, there is general agreement that this
similarity
is not based on common ancestry and most classification systems place
this
family in a position well removed from families of the Hamamelidae.
This points out the importance of the gynoecium in that the primary
structural
difference between this family and the other 'amentiferous'
flowering plants is the capsular fruit. As indicated by
the
local name for our most common poplar - 'cottonwood' - seeds emerging
from
capsules of the Salicaceae are comose
(with a tuft of trichomes) and, as a result, all reproductive events -
pollination, fertilization and dispersal of the product - are
wind-mediated
in this family.
The single representative of Salix
in our local flora - S. nigra:
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| Staminate catkin at anthesis | Pistillate catkin at anthesis | Pistillate catkin with fruit and comose seeds |
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| Staminate catkin of Salix with dark bract subtending each flower | Populus nigra - overview - from Kohler's Medicinal Plants |
The aspen (Populus
tremuloides)
is a rhizomatous
woody perennial. A single plant can therefore produce many shoots
and a 'stand' of aspen often represents a single plant.
The relatively recent capability to employ molecular markers to
identify
individuals has allowed large 'clones' to be defined and, given the
growth
rate for this plants, some of the larger aspen clones might be the
oldest
living individuals.
More information on the Salicaceae