The Asteridae
Family Overview - Lamiales

Verbenaceae - the Vervain or Verbena Family

Diversity:  Shrubs, trees, and herbs with the latter predominating in our flora -  about 100 genera and 2, 600 species.

Distribution:  Worldwide, but as opposed to other elements of the Lamiales, with greatest diversity in the tropics. We have 17 genera and 64 species in the Texas flora

Floral structure:

Significant features:  This family carries the distinctive gynoecium of the Lamiales in that the ovary usually shows a second septum, producing 4 locules and each of these has only a single seed.  However, the lobing in most taxa is not as extensive as that found in the Boraginaceae and Lamiaceae and the style is not gynobasic, it is usually positioned at the tip of the ovary.  Taxa of the Verbenaceae, especially the herbs, tend of show quadrangular stems and all have opposite leaves.  This similarity to the Lamiaceae also extends to corolla zygomorphy but this is less developed in the Verbenaceae and usually expressed as an asymmetry in the corolla lobes as opposed to the bilabiate ('two-lipped') corolla configuration common among the mints..

Lantana includes of ca. 150 shrubby species, mostly from the American tropics.  Our flora includes native wild species and 'escapes' from cultivation.  Note weak zygomorphy (one petal lobe larger than the others) characteristic of the family


ornamental hybrid
racemose inflorescence
capitate inflorescence

Callicarpa americana:

at anthesis
just past anthesis
fruit - mid summer
mature fruit

More information on the Verbenaceae

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